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Strümpher, Werner P, Sole, Catherine L, Villet, Martin H, Scholtz, Clarke H. Brill; 2016. Allopatric speciation in the flightless Phoberus capensis (Coleoptera: Trogidae) group, with description of two new species.
Strümpher, Werner P, Villet, Martin H, Sole, Catherine L, Scholtz, Clarke H. Brill; 2016. Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera Scarabaeoidea)..
Strümpher, Werner P, Sole, Catherine L, Villet, Martin H, Scholtz, Clarke H. Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. 2014.